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Dear all,

If you know of any promising UK/EU students looking for a PhD 
studentship in bird or insect flight biomechanics, please could you pass 
on to them the following advertisement for three fully-funded 
studentships in the Oxford Animal Flight Group. 

Details also at: 
http://users.ox.ac.uk/~zool0261/vacancies.html

All three projects have biomimetic applications relevant to Unmanned and Micro Air Vehicle control control systems and design.

best wishes,
Richard Bomphrey



Three fully funded studentships available in the Oxford Animal Flight 
Group: deadline 21 January 2011

We are pleased to be able to offer up to three fully funded PhD 
studentships this year. Please scroll down to see the full advertisement 
for each of the three studentships. Please note that the eligibility 
criteria and starting dates differ between the three studentships.

1. Vision-based flight control in insects, with applications to unmanned 
air vehicles

Supervisors: Dr Graham Taylor; Dr Holger Krapp

Feedback control is essential to the flight of insects and most modern 
aircraft, but there the resemblance ends. Aircraft control systems 
employ a few high-quality sensors to measure or estimate their current 
state with a high degree of accuracy; insects have numerous sensors of 
lower quality, which sense changes in state rather than absolute state. 
Aircraft control systems typically have their sensors arranged 
orthogonally; insects have their sensors arranged non-orthogonally. 
Aircraft control systems process state feedback centrally; insects 
process signals at the periphery of their nervous system and combine 
input from different sensory modalities before using this as feedback. 
In light of these differences, it has recently been suggested that the 
sensory systems of insects are configured so that they are specifically 
tuned to detect excitation of their rigid body modes of motion (Taylor & 
Krapp, 2007). The aim of this studentship is to investigate the 
mode-sensing hypothesis, by using a combination of theoretical and 
experimental approaches, so as to inform the design of control systems 
for unmanned air vehicles (UAVs). The hypothesised control architecture 
is fundamentally different from the architectures employed in aircraft 
control systems, and offers the potential for exploiting noisy and datum 
deficient measurements in the feedback control of a Mini/Nano UAV. This 
could include, for example, input from sensors measuring optic flow.

The studentship will be held in the Oxford Animal Flight Group, 
Department of Zoology, Oxford University (supervisor: Dr Graham Taylor), 
but is being run in collaboration with the Insect Neurophysiology Group, 
Department of Bioengineering, Imperial College (co-supervisor: Dr Holger 
Krapp). The ideal candidate would EITHER have a background in 
Engineering or Applied Mathematics and an interest in thinking out of 
the box, OR would have a background in Biological Sciences and a 
willingness to think mathematically.

To apply please use the online application system at: 
http://www.ox.ac.uk/admissions/postgraduate_courses/apply/ . Please 
remember to quote the studentship reference code GT01. Because you will 
be applying for a pre-defined project, please provide a Statement of 
Purpose, and not a Research Proposal, at the relevant point in the 
application. Please use this statement to explain your specific interest 
in the advertised project. You may, if you wish, incorporate material 
from this advertisement. Any queries regarding the application procedure 
please contact [log in to unmask] The closing date is Friday 
21st January 2011. The successful candidate will start their studentship 
as soon as possible after 13 March 2011, and in any case before the end 
of September 2011. The University of Oxford is an Equal Opportunities 
employer.

Funding Notes
This is an Industrial CASE studentship, which is being jointly funded by 
EPSRC and Dstl (Defence Science and Technology Laboratory). In order to 
assist recruitment to this position, the industrial component of the 
funding will be used to elevate the stipend by £1500 per annum over the 
national minimum stipend rate for Research Council studentships. 
Eligibility for this studentship is limited to UK nationals only. The 
full award is available to UK nationals who meet the UK residency 
requirements, while UK students who do not meet the UK residency 
requirements are eligible for a studentship covering tuition fees. 
Further information can be found at: 
http://www.epsrc.ac.uk/funding/students/Pages/eligibility.aspx

Taylor, G. K. & Krapp, H. G. (2007). Sensory systems and flight 
stability: what do insects measure, and why? Adv. Insect Physiol., 34, 
231-316. doi:10.1016/S0065-2806(07)34005-8


2. Avian flight mechanics: atmospheric energy extraction and gust 
response in birds

Supervisors: Dr Graham Taylor; Prof. Adrian Thomas

Birds are masters of their aerial environment. One of their most 
impressive accomplishments is their ability to deal with – and even 
exploit – atmospheric heterogeneity at small spatial scales. Birds are 
able to negotiate the gustiest of conditions, to thermal-soar on days 
when a sailplane could not leave the ground, and to ascend vertically 
over a cliff edge without once flapping their wings. The mechanistic 
detail of how birds accomplish these feats is largely unknown, but 
similar sensory, aeroelastic, and muscular mechanisms are likely to be 
common to all. The aim of this studentship is to make detailed 
measurements of the flight dynamics and flight performance of trained 
birds of prey using onboard inertial instrumentation and high-speed 
photogrammetry, and to analyse these in combination with information 
from onboard pressure sensors and ultrasonic anemometers to understand 
how birds deal with and exploit atmospheric heterogeneity. The project 
will focus upon three specific behaviours: response to mechanical and 
thermal turbulence, exploitation of thermals in cross-country soaring, 
and exploitation of wind shear over sea cliffs. The project is intended 
foremost to be a study of the flight mechanics of birds, but there are 
obvious engineering applications to the flight of Unmanned Air Vehicles, 
for which atmospheric energy extraction and gust response properties 
modelled on birds would be of great commercial and strategic significance.

To apply please use the online application system at: 
http://www.ox.ac.uk/admissions/postgraduate_courses/apply/ . Please 
remember to quote the studentship reference code DTG6. Because you will 
be applying for a pre-defined project, please provide a Statement of 
Purpose, and not a Research Proposal, at the relevant point in the 
application. Please use this statement to explain your specific interest 
in the advertised project. You may, if you wish, incorporate material 
from this advertisement. Any queries regarding the application procedure 
please contact [log in to unmask] The closing date is Friday 
21st January 2011. The successful candidate will start their studentship 
in October 2011. The University of Oxford is an Equal Opportunities 
employer.

Funding Notes:
UK Research Council Competition Funded Studentship. These awards are 
available to UK nationals and EU students who meet the UK residency 
requirements, while EU students who do not meet the UK residency 
requirements are eligible for studentships covering tuition fees. 
Further information can be found at:
http://www.bbsrc.ac.uk/funding/studentships/studentship_eligibility.pdf

Carruthers, A. C., Thomas, A. L. R. & Taylor, G. K. (2007). Automatic 
aeroelastic devices in the wings of a Steppe Eagle Aquila nipalensis. J. 
Exp. Biol. 210, 4136-4149. doi:10.1242/jeb.011197

Carruthers, A. C., Walker, S. M., Thomas, A. L. R. & Taylor, G. K. 
(2010). Aerodynamics of aerofoil sections measured on a free-flying 
bird. Proc. Inst. Mech. Eng. Part G - J. Aerosp. Eng. 224, 855-864. 
doi:10.1243/09544100JAERO737


3. Wing shape, kinematics and performance in insect flight

Supervisors: Dr Richard Bomphrey; Prof. Adrian Thomas

This inter-disciplinary project analyses the fundamental features of 
insect wing shape and motion that are important for specific ecological 
tasks. Research output will constitute the most comprehensive study to 
date of how the apparatus for flight manipulates, and is constrained by, 
the physical world. The student will follow two themes to explore the 
relationship between wing kinematics and flight performance. They will 
use (1) high-speed cameras to record detailed topographical wing 
kinematics from a range of insects chosen for their contrasting ecology, 
and (2) artificial selection to examine the effects of alternative 
morphological parameters on performance.

Kinematics: The student will record detailed wing topographies 
throughout the wingbeat cycle for representative species. Wing shape and 
kinematic data will then be used to identify the morphological and 
kinematic variables which are critical for certain ecological roles. 
This will be achieved using an existing state-of-the-art photogrammetric 
technique developed within the Flight Group. Tests will begin by 
quantifying variation across wingbeats within individuals, before 
progressing to quantify intra- and inter-specific differences. Finally, 
the student will integrate their experimental design with an 
EPSRC-funded Postdoc so that aerodynamic output is simultaneously 
recorded from experimental subjects.

Artificial selection: The student will select strains of fruit fly for 
exaggerated wing parameters (larger area, higher aspect ratio) and 
measure consequent changes in performance (speed, load lifting, aerial 
agility) within a flight arena. The inverse of the problem will be run 
in parallel, i.e. identification of salient performance criteria and 
analysis of the kinematic and morphological properties that make them 
possible.

Future direction: This work naturally leads to further collaborative 
Computational Fluid Dynamics projects (e.g. Young, et al. Science 2009), 
and Finite Element modelling projects that could further inform the 
design of miniature vehicles. This PhD could also form the foundation 
for further work in eco-morphology.

To apply please use the online application system at: 
http://www.ox.ac.uk/admissions/postgraduate_courses/apply/ . Please 
remember to quote the studentship reference code DTG8. Any queries 
regarding the application procedure please contact 
[log in to unmask] The closing date is Friday 21st January 
2011. The successful candidate will start their studentship in October 
2011. The University of Oxford is an Equal Opportunities employer.

Funding Notes:
UK Research Council Competition Funded Studentship. These awards are 
available to UK nationals and EU students who meet the UK residency 
requirements, while EU students who do not meet the UK residency 
requirements are eligible for studentships covering tuition fees. 
Further information can be found at:
http://www.bbsrc.ac.uk/funding/studentships/studentship_eligibility.pdf
http://www.nerc.ac.uk/funding/studentships/studentship_eligibility.pdf

__________________________________________
Dr Graham K. Taylor
Animal Behaviour Research Group
Department of Zoology, Oxford University
Tinbergen Building, South Parks Road
Oxford, OX1 3PS
United Kingdom
tel. +44 (0)1865 271219
fax. +44 (0)1865 310447
http://users.ox.ac.uk/~zool0261

__________________________________________
Dr Richard J. Bomphrey
EPSRC CAF
Animal Behaviour Research Group
Department of Zoology, Oxford University
Tinbergen Building, South Parks Road
Oxford, OX1 3PS
United Kingdom
tel. +44 (0)1865 271224
fax. +44 (0)1865 310447