I have had a number of responses to my query - thank you all, it is much appreciated and (pleasingly!) confirms my original thoughts on this. For anyone who's interested - my confusion arose when reading the section on page 5 of this guidance document:
http://www.nationalarchives.gov.uk/documents/information-management/copyright-related-rights.pdf
It states that
"the first owner(s) of copyright will be:...
the owner of the negative - in the case of a photograph taken between 1 July 1912 and 31 July 1989"
The guide goes on to say that:
"The present owner of any of these copyrights will often, but not always, be the direct descendant(s) or successor (such as a successor company) of the first owner."
This is where my confusion arose - I had originally assumed that copyright was with the collection's heirs, but as this stated that original copyright ownership was with the owner of the physical object (specifically negatives) this appeared different to other copyright examples I had come up again.
I appear to have overcomplicated this issue - once again, thank you for all the responses.
Best wishes
Kelda
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